Nathan Fillion is a Canadian actor best known for his portrayal of the lead role of Captain Malcolm Reynolds in the television series Firefly and its feature film continuation, Serenity, as well as the role of Richard Castle on the ABC series Castle. He has acted in traditionally distributed films like Slither, Internet-distributed films like Dr. Horrible’s Sing-Along Blog, television soap operas and sitcoms, and theater, and voice acted in the video games Halo 3, Halo 3: ODST and Halo: Reach as well as Wonder Woman, Green Lantern: Emerald Knights, Justice League: Doom and Monsters University.
He was cast as James Frederick “The Minnesota” Ryan in Steven Spielberg’s Saving Private Ryan. Fillion had a recurring role as Caleb in 2003 in the final five episodes of the final season of Joss Whedon’s series Buffy the Vampire Slayer. In 2002, Fillion starred as Captain Malcolm Reynolds in the Joss Whedon science fiction television series Firefly. Fillion called his time on Firefly the best acting job he ever had, and compares every job he has had to it. Although the show was cancelled, it was adapted to the big screen; he reprised his role as Mal in Whedon’s movie Serenity (2005).
Fillion lent his voice to the animated series King of the Hill in 2001, the video game Jade Empire (as the voice of Gao the Lesser), and the animated series Justice League Unlimited(as Vigilante in the episodes “Hunter’s Moon” and “Patriot Act”) in 2005 and 2006. Fillion starred in James Gunn’s (Guardians of the Galaxy) 2006 horror film Slither.
Fillion starred in the romantic comedy film Waitress, in horror sequel White Noise 2: The Light, and LOST, as Kevin, Kate’s ex-husband. Fillion was cast as Alex Tully in the series Drive, and joined the cast of ABC’s Desperate Housewives as Dr. Adam Mayfair. Fillion stars as the title character Richard Castle, a mystery novelist who helps the NYPD solve crimes in ABC television series Castle. He recently appeared as Hermes in the YA adaptation film Percy Jackson: Sea of Monsters.